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Catalogue of the ungulate mammals in the British Museum (Natural History) (Volume 5) online

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UC-NRLF




i !



.







WILLIAM DILLER MATTHEW




MA

m/^



PRESENTED



BY



The Trustees



OF



THE BRITISH MUSEUM



B.M (N H.) CAT. UNO. MAMM. V.




Photo, by Maull and Fox.

RICHARD LYDEKKER, F.R.S,



CATALOGUE

OP THE

UNGULATE MAMMALS

IN THE

BRITISH MUSEUM (NATURAL HISTORY).

VOL. V.
PERISSODACTYLA

(HOKSES, TAPIKS, BHINOCEROSES),

HYRACOIDEA

(HYRAXES),

PROBOSCIDEA

(ELEPHANTS).



WITH ADDENDA TO THE EARLIER VOLUMES.



BY THE LATE RICHARD LYDEKKER, F.R.S.



LONDON: ?ji*

PRINTED BY ORDER OF THEf TRUSTEED OT .TJJ0S.



BRITISH MUSEUM.
SOLD BY

LONGMANS, GEEEN & Co., 39, PATERNOSTER Row, LONDON, E.G.

B. QUARITCH, 11, GRAFTON STREET, NEW BOND STREET, LONDON, W.

DULAU & Co., LTD., 37, SOHO SQUARE, LONDON, W.

AND AT THE

BRITISH MUSEUM (NATURAL HISTORY), CROMWELL ROAD, S.W.
1916.

(All rights reserved.)




EARTH

SCIENCE*

UBRARY



LONDON :

PRINTED BY WILLIAM CLOWES AND SON'S, LIMITED,
DUKE STREET, STAMFORD STREET, S.E., AND GREAT WINDMILL STREET, W.



PREFACE

THE hope was expressed, in the Preface to the previous
Volume, that it might be possible to issue a fifth, or
concluding Volume, without undue delay. That hope has
been realised, as the result of finding the materials for the
completion of the work among the papers of the late
Mr. Richard Lydekker.

The volume now published is practically as the author
left it, and the only alterations that have been made are
verbal ones. Even the illustrations are those which he had
himself selected.

It will be obvious, however, that this volume should not
be regarded as a finished production. The earlier part is
drawn up on the general lines of the preceding volumes,
and contains lists of the specimens in the collection. These
do not appear in the second half of the volume, which
consists mainly of a list of species and subspecies, with
synonymy and indications of the typical localities. The
manuscript was, indeed, in the condition of a preliminary
draft, and renders should bear in mind that it has not had
the advantage of being amplified and revised by the author.

On the other hand, it has not been thought advisable to
make any attempt to complete the volume on the full
original plan, partly on account of the difficulty of finding
anyone competent to do so, and partly because the mixed
responsibility for the work which would be thereby intro-
duced would not be for the benefit of science.

The thanks of the Trustees are due to Messrs. George
Allen & Unwin, Ltd., for permission to use certain copyright
figures published in Mr. Lydekker's work, The Horse and its
Relatives; to Lord Rothschild, F.R.S., for the use of figures
from the Novitates Zoological ; and to the Zoological Society

808133



IV PREFACE

of London, the United States National Museum, and Major
Stanley S. Flower for other figures, as indicated in the
respective legends. Their thanks are also due to Dr. J.
Biittikofer and to Mr. F. A. 0. Pyni for photographs which
have served as the originals of other figures, and to the
Duchess of Bedford and to Mr. E, I. Pocock, F.K.S., for
information about certain specimens.

SIDNEY F. HAPvMEE,

Keeper of Zoology.

BRITISH MUSEUM (NATURAL HISTORY),
LONDON, S.W.

January 26^, 1916.



CONTENTS

PAGE

PREFACE .......... iii

SYSTEMATIC INDEX TO VOL. Y. v

LIST OF ILLUSTRATIONS ... . . x iii

SYSTEMATIC INDEX TO VOLS. I.-V. xv

ORDER UNGULATA.

SUBORDER II. PERISSODACTYLA ...... 1

FAMILY I. EQUID.E 3

GENUS EQUUS . ..... 4

1. SUBGENUS EQUUS ....... 6

I. EQUUS CABALLUS ....... 6

A. Eqmis caballus caballus .... 6

B. Eqmis caballus przewalskii .... 7

II. EQUUS KIANG ....... 8

III. EQUUS HEMIONUS ...... 11

IV. EQUUS ONAGER ....... 12

A. Equus onager indicus ..... 12

B. Equus onager castaneus . . . .13

C. Equus onager henrippus .... 14

D. Equus onager onager . . . . .14

2. SUBGENUS DOLICHOHIPPUS ..... 15

V. EQUUS (DOLICHOHIPPUS) GREVYI .... 17

A. Equus grevyi grevyi ..... 18

B. Equus grevyi berberensis .... 18

3. SUBGENUS HIPPOTIGRIS ...... 19

VI. EQUUS (HlPPOTIGRIS) QUAGGA .... 19

A. Equus quagga quagga ..... 20

B. Equus quagga burchelli .... 21

C. Equus quagga antiquorum .... 22
D. Equus quagga pococki .... 23
E. Equus quagga chapman! .... 23
F. Equus quagga wahlbergi .... 24



VI CONTENTS

PAGK

G. Equus quagga transvaalensis . . .25
H. Equus quagga selousi . * ... 25
I. Equus quagga boehmi . ... . 26

J. Equus quagga zambesiensis . . .26
K. Equus quagga crawshayi .... 27
L. Equus quagga cuninghamei ... 28

M. Equus quagga grant! . . . . 80

VI F. EQUUS (HIPPO-TIGRIS) FOAI .... 32

VIII. EQUUS (HIPPOTIGRIS) ZEBRA . .. . .34

A. Equus zebra zebra . . . . .35
B. Equus zebra hartmannae . fc . .36

4. SUBGENUS ASINUS . . . . . . .36

IX. EQUUS (AsiNus) ASINUS . . . . 36
A. Equus asinus asinus . . . .37
B. Equus asinus africanus . . . .37
C. Equus asinus taeniopus . , . .38
D. Equus asinus somaliensis . . v . . 38

FAMILY II. TAPIRID^ . . . . . . .39

GENUS TAPIRUS . . . . . . . . 39

1. SUBGENUS ACROCODIA . . . . . . 39

I. TAPIRUS (ACROCODIA) INDICUS . . . . 39

2. SUBGENUS TAPIRUS . . . . . . . 41

II. TAPIRUS TERRESTRIS . . . ... .41

A. Tapirus terrestris terrestris . . . . 42

B. Tapirus terrestris laurillardi . .' . 43

C. Tapirus terrestris mexianse . . . .43

D. Tapirus terrestris spegazzinii . . . 43

III. TAPIRUS ROULINI . . . .44

3. SUBGENUS TAPIRELLA . . . . . .44

IV. TAPIRUS (TAPIRELLA) BAIRDI . . . .45

V. TAPIRUS (TAPIRELLA) DOWI . . . . . 46

FAMILY III. BHINOCEROTIDJE . . . . . .46

GENUS EHINOCEROS. . . , , . - . . 46

1. SUBGENUS RHINOCEROS . , . . .46

I. RHINOCEROS UNICORNIS ...... 47

II. RHINOCEROS SONDAICUS ..... 48

2. SUBGENUS DICERORHINUS ...... 49

III. RHINOCEROS (DICERORHINUS) SUMATRENSIS . . 49
A. Rhinoceros sumatrensis sumatrensis . . 50
B. Rhinoceros sumatrensis blythi . . , 50
C. Rhinoceros sumatrensis lasiotis 51



CONTENTS VJi

1'AI.K

3. SUBGENUS DlCEROS ....... 51

IV. RHINOCEROS (DlCEROS) BICOKMs .... 52

A. Rhinoceros bicornis bicornis . . . 54

B. Ehinoceros bicornis holmwoodi ... 54

C. Rhinoceros bicornis somaliensis . . . 55

D. Rhinoceros bicornis cucullatus . . . 5.">

V. RHINOCEROS (DICEROS) SIMUS .... 55

A. Rhinoceros simus simus .... 50

B. Rhinoceros simus cottoui .... 56

SUBORDER III. HYRACOIDKA ....... 58

FAMILY PROCAVIIDJS ........ 58

GEN 7 us PROCAVIA ........ 58

1. SUBGENUS PROCAVIA ...... 58

I. PROCAVIA CAPENSIS ...... 58

A. Procavia capensis capensis .... 60

B. Procavia capensis windhuki .... 60

C. Procavia capensis reuningi .... 60

D. Procavia capensis marlothi .... 60

E. Procavia capensis schultzei .... 61

F. Procavia capensis waterbergensis ... 61

II. PROCAVIA NIGRICANS .... .61

III. PROCAVIA JOHNSTONI . . . .61

IV. PROCAVIA SCIOANA ...... 62

V. PROCAVIA ERLANGERI ...... 62

VI. PROCAVIA ALPINI ...... 62

VII. PROCAVIA KERSTINGI ...... 62

VIII. PROCAVIA GOSLINGI ...... 62

IX. PROCAVIA SYRIACA ...... f>;5

A. Procavia syriaca syriaca .... 64

B. Procavia syriaca jayakari .... 64

X. PROCAVIA RUFICEPS ...... 64

XI. PROCAVIA BURTONI ...... 64

XII. PROCAVIA SHARICA ... 65*

XIII. PROCAVIA HABESSINICA ..... (;.">
A. Procavia habessinica habessinica 66
B. Procavia habessinica minor .... 66

XIV. PROCAVIA FERRUGINEA ..... 66

XV. PROCAVIA PALLIDA .... 66

XVI. PROCAVIA RUWENZORII . 66



Vlll CONTENTS

PAGE

XVII. PROCAVIA MACKINDERI ..... 67
A. Procavia mackinderi mackinderi ... 67
B. Procavia mackinderi zelotes .... 67

XVIII. PROCAVIA LOPESI . . ' . . . 67

XIX. PROCAVIA OWENI . . ... .67

XX. PROCAVIA JACKSONI . . . . f . 68

XXI. PROCAVIA D^MON .. . ... . 68

XXII. PROCAVIA MATSCHIEI . .. ... . . 68

XXIII. PROCAVIA WELWITSCHI . . . . .68

XXIV. PROCAVIA VOLKMANNI . . . . .69

XXV. PROCAVIA LATASTEI . . . . . .69

XXVI. PROCAVIA BRUCEI . . ... .69

A. Procavia brucei brucei . . .69

B. Procavia brucei somalica . . . . 70

C. Procavia brucei hindei . . ... 70

D. Procavia brucei borana . . . . 70

E. Procavia brucei ruddi . . . . .71

F. Procavia brucei maculata . . .71

G. Procavia brucei grant! . ." . . 71

H. Procavia brucei manningi . . . .72

I. Procavia brucei thomasi . . . .72

J. Procavia brucei kempi ' . t . ; . 72

K. Procavia brucei princepa . . , . 73

XXVII. PROCAVIA PUMILA . . . .73
A. Procavia pumila pumila . . . > . 73
B. Procavia pumila rudolfi . 73

XXVIII. PROCAVIA SLATINI . . . .74

XXIX. PROCAVIA MUENZERI . . . . .74

XXX. PROCAVIA FROMMI . . . . . .74

XXXI. PROCAVIA SCHUBOTZI . . ... .74

XXXII. PROCAVIA BOUNHIOLI . . . . . 74

XXXIII. PROCAVIA BUTLERI . . . .74

XXXIV. PROCAVIA BAMEND.E . ... .; . 75

XXXV. PROCAVIA KAMERUNENSIS .... 75
2. SUBGENUS DENDROHYRAX ...... 75

XXXVI. PROCAVIA (DENDROHYRAX) SCHEELEI . . 75

XXXVII. PROCAVIA (DENDROHYRAX) VALIDA . . 75

XXXVIII. PROCAVIA (DENDROHYRAX) EMINI . . 76
A. Procavia emini emini . . . . 76
B. Procavia emini latrator . ....'- . . 76^



CONTENTS ix

XXXIX. PROCAVIA (DENDROHYRAX) ARBOREA . . 70

XL. PROCAVIA (DENDROHYRAX) STUHLMANNI 77

XLT. PROCAVIA (DENDROHYRAX) NEUMANNI ... 77

XLII. PROCAVIA (DENDROHYRAX) DORSALIS . 77

XLIII. PROCAVIA (DENDROHYRAX) MARMOTA . . 78

XLIV. PROCAVIA (DENDROHYRAX) BETTONI ... 78

XLV. PROCAVIA (DENDROHYRAX) ASCHANTIENSIS . . 78

XLVI. PROCAVIA (DENDROHYRAX) TERRICOLA . . 78

XLVII. PROCAVIA (DENDROHYRAX) ADAMETZI . . 78

A. Procavia aclametzi adametzi . . 78

B. Procavia adametzi zenkeri .... 79

XLVIII. PROCAVIA (DENDROHYRAX) TESSMANNI . . 79

XLIX. PROCAVIA (DENDROHYRAX) CRAWSHAYI . . 79

A. Procavia crawshayi crawshayi ... 79

B. Procavia crawshayi laikipia . ... 79

L. PROCAVIA (DENDROHYRAX) ADOLFI-FRIEDERICI . . 79

LI. PROCAVIA (DENDROHYRAX) SCHEFFLERI ... 80

SUBORDER IV. PROBOSCIDEA ..... 80

FAMILY ELEPHANTIDJE ..... .80

GENUS ELEPHAS ........ 80

1. SUBGENUS ELEPHAS . ... 81

I. ELEPHAS MAXIM us ...... 81

A. Elephas maximus maxiinus .... 82

B. Elephas maximus zeylanicus ... 83

C. Elephas maximus sumatranus ... 83

D. Elephas maximus hirsutus .... 84

2. SUBGENUS LOXODONTA ...... 85

II. ELEPHAS (LOXODONTA) AFRICANUS ... 85
A. Elephas africanus capensis .... 86
B. Elephas africanus selousi . . 87

C. Elephas africanus cyclotis .... 87

D. Elephas africanus cottoni .... 87
E. Elephas africanus knochenhaueri . . 88
F. Elephas africanus peeli .... 88
G. Elephas africanus cavendishi 89
H. Elephas africanus oxyotis .... 89
I. Elephas africanus orleansi .... 90
J. Elephas africanus rothschildi . . .91
K. Elephas africanus albertensis ... 91



LIST OF ILLUSTBATIONS

PAGE

Fig. 1. Bones of the Left Fore-foot of a Horse (A) and of a

Rhinoceros (B) ....... 2

2. Skull of Female Tarpan, or Mongolian Wild Horse

(Equus caballus przewalskii) .... 4

3. The Eight Upper Cheek-teeth of the Tarpan, or
Mongolian Wild Horse (Equus caballus przewal-
sJcii). (From Lydekker, The Horse and its Rela-
tives, 1912) 5

,, 4. Female Tarpan, or Mongolian Wild Horse (Equus

caballus przeiualsltii) ...... 8

,, 5. Female Kiang (Equus Jciang}. (From Lydekker, Proc.

Zool. Soc. 1904) 10

,, 6. Female Kobdo Onager (Equus onager castaneus}.

(From Lydekker, Novitates Zoologicx, 1904) . 13

,, 7. Head of Grevy's Zebra (Equus [Dolichohippus']
grevyi). (From Lydekker, The Horse and its Rela-
tives, 1912) . . 15

,, 8. Head of Grevy's Zebra (Equus [Dolichohippus]

grevyi) ........ 16

,, 9. Grevy's Zebra (Equus {Dolichohippus'] grevyi) . . 17

,, 10. The True, or Cape Quagga (Equus quagga quag g a] . 20
,, 11. Cuninghame's Zebra (Equus quagga cuninghamei) .

(From a photograph by Mr. F. A. 0. Pym) . . 28

,. 12. Cuninghame's Zebra (Equus quagga cuninghamei).

(From a photograph by Mr. F. A. 0. Pym) . . 29

,, 13. Male Masai Zebra (Equus quagga granti). (From

Sclater, Proc. Zool. Soc. 1901) .... 31

,, 14. Skin of Foa's Zebra (Equus [Hippotigris] foai).

(From Lydekker, The Horse and its Relatives, 1912) 33

,, 15. The Mountain Zebra (Equus {Hippotigris} zebra] . 34

,, 16. Male Nubian Wild Ass (Equus asinus africanus}.

(From Lydekker, Novitates Zoologicx, 1904) . . 37
,, 17. Head of a Great Indian Ehinoceros (Rhinoceros uni-

cornis) . . . . . . . .47



XIV LIST OF ILLUSTRATIONS

PAGE

Fig. 18. Heads of (a) White Rhinoceros (Rhinoceros [Diceros]

simus), and (b) Black Rhinoceros (R. [D.J bicornis) 52

,, 19. Upper Aspect of Skull of A, Rhinoceros bicornis, and
B, R. bicornis somaliensis. (From Lydekker,
Proc. Zool. Soc. 1911) ...... 53

,, 20. Male Nile White Rhinoceros (Rhinoceros simus cot-

toni). (From Heller, Smithson. Misc. Coll. 1913) . 57

,, 21. The Syrian Hyrax (Procavia syriaca) . . . 63

,, 22. An Upper Molar (A) of the African Elephant
(E. africanus), and (B) of the Asiatic Elephant
(Elephas maximus) . . . .... . 80

? 23. Large Male Indian Elephant (Elephas maximus).

(From S. S. Flower, "Report on a Zoological

Mission to India in 1913," Ministry of Public Works,

Cairo, 1914) ~ 4 . , . . . . 81
., 24. Male Elephant (Elephas maximus sumatranus).

(From a photograph sent by Dr. J. Biittikofer) . 83
,, 25. Young Female Malay Elephant (Eleplias maximus

hirsutus) . . . . \. . . . 84
,, 26. Skull of African Elephant (ElepTias africanus) . . 85
,, 27. Head of a Female Cape Elephant (ElepTias africanus

capensis). (From Lydekker, Proc. Zool. Soc. 1907) 86
,, 28. Head of Male Elephant (Elephas africanus Ttnochen-

haueri ?). (From Lydekker, Proc. Zool. Soc. 1907) . 88

,, 29. Head of an Abyssinian Elephant (Elephas africanus

oxyotis). (From Lydekker, Proc. Zool. Soc. 1907) . 89

,, 30. Skull of Abyssinian Elephant (Elephas africanus

oxyotis). (From Lydekker, Proc. Zool. Soc. 1907) . 90

31. Skull of Semliki Elephant (Elephas africanus alber-

tensis). (From Lydekker, Proc. Zool. Soc. 1907) . 91



GENERAL SYSTEMATIC INDEX.

(VOLS. i.-v.)



Order UNGULATA.

Suborder I. ARTIODACTYLA.

Section A. PECORA.

Family I. BOVID^E.

Subfamily i. BOVINE.

Genus Bos.

1. Subgenus Bos. . L

I. Bos taurus ........ 12

2. Subgenus BIBOS.

II. Bos (Bibos) gaurus 13

A. Bos gaurus gaurus ...... 18

B. ,, readi 20

C. ,, ,, hubbacki ...... 20

III. Bos (Bibos) banteng . . .21

A. Bos banteng banteng ...... 23

B. lowi 25

C. .. birmanicus ..... 27
D porteri 28

E. .. ,, butleri 29

F. ,, ,, subsp. ...... 29

3. Subgenus POKPHAGUS.

IV. Bos (Poephagus) grunniens ..... 31

,, grunniens mutus ...... 33

4. Subgenus BISON.

V. Bos (Bison) bonasus ...... 35

A. Bos bonasus bonasus ...... 36

B. ,, ,, caucasius ..... 36

VI. Bos (Bison) bison ... ... 37

A. Bos bison bison . . .... 38

B. ,, athabascse . ... 39
V. b



XVI



GENERAL SYSTEMATIC INDEX



B.
C.
D.



5. Subgenus BUBALUS.

VII. Bos (Bubalus) bubalis . . , '

A. Bos bubalis bubalis

B. ,, ., macroceros

C. fulvus

D. ,, hosei

VIII. Bos (Bubalus) mindorensis , .

IX. Bos (Bubalus) depressicornis .

A. Bos depressicornis depressicornis

B. ,, ,, fergusoni . .

X. Bos (Bubalus) caffer

A. Bos caffer caffer . V
neumanni
ruahensis.

schillings! , . ,
athiensis . . '" .
wemberensis
gariepensis
radcliffei .
limpopoensis
azrakensis

wiesei . . .
aequinoctialis
mathewsi .
cottoni
thierryi

mayi . . . .

brachyceros
houyi . - .
planiceros .
adamause .
hunti
nanus
beddingtoni
diehli
hyleeus
adametzi .



E.

F.

G.

H.

I.

J.

K.

L.

M.

N.

O.

P.

P 1 .

Q.

Q 1 -

K.

S.

T.

T 1 .

T 2 .

T 3 .

T 4 .

U.



nuni
simpsoni



Vol. I.


V.


PAGE


PAGE


. 41




. 43




. 45




45




. 46




. 47




. 48




. 49




. 50




. 50




. 52




. 53




. 53




54




.


92


. 54




55




. 55




. 56




57




. 57




. 58


94


59




. 60




. 61




. 62




. 62


94




94


! 66




.


95


. 67


95


. 68




. 70






95


.


96


. .


96


. .


97



Other names applied to buffaloes from British and
German East Africa . . -. ' .

Subfamily ii. CAPBIN.E.
I. Genus Ovis.

I. Ovis aries ......

II. Ovis musimon * . ,

III. Ovis orientalis . . . .

A. Ovis orientalis orientalis . . . '- ,

B. ,, gmelini . .

C. ,, ,, anatolica . . .

D. ,, urmiana

E. ,, ,, erskinei .....

F. ,. M isphahanica . .



70



75

75 97

77
79
80
81
81
82
83



GENERAL SYSTEMATIC INDEX



XVll



IV. Ovis laristanica



Vol. I. V.
PAGE PAGE



V. Ovis vignei ..... 84

A. Ovis vignei vignei 85

B. ,, ,, punjabiensis . 86

C. ,, ., cycloceros ..... 88

D. arkar .... 90

VI. Ovis ammon ..... 92

A. Ovis ammon ammou ..... 95
A 1 . ,, , kozlovi ..... 97
, mongolica ... .96

C. , hodgsoni ..... 97

D. , , storcki .... 100

E. , sairensis . . . 101

F. , littledalei .....' 102

G. , nigrimontana .... 103 -98

G 1 . , severtzovi . . . . ... 98

H. , karelini . . 104

I; , fceinsi - . 105

1 . , collmm . . . . . ... 98

J. , humei . . 105

K. , poli 106

Other races described ...... 109 99

VII. Ovis canadensis ....... 109

A. Ovis canadensis canadensis . . . .111

B. , ,, californiana .... 113

C. , ,, auduboni . . . . .113

D. , ,, nelsoni ..... 114

E. . sieme 115

F. , ,, cremnobates .... 115

G. , mexicana ..... 116

H. , gaillardi 116

I. i M dalli 118

J. , kenaiensis . . . . . 119

K. , ,, fannini ..... 119

L. stonei 120

M. cowani 120

N. ,, ,, nivicola .... 121

O. borealis 121 99

P. aUeni . . . 122

II. Genus AMMOTRAGUS.

Ammotragus lervia . . . . . . .123

A. Ammotragus lervia lervia ..... 124

B. sahariensis .... 125

C. ,, ornata 125

D. ,, ,, blainei .... 126



Pseudoi's nahoor



III. Genus P s E u D o i' s.



. 126
b 2



XV111



GENERAL SYSTEMATIC IXDEX



IV. Genus CAPRA.

I. Capra caucasica ....

A. Capra caucasica cylindricornis

B. ,, ,, caucasica .

II. Capra severtzowi ....

A. Capra severtzowi severtzowi

B. .. ,, dinniki
Incertae Sedis .

III. Capra pyrenaica . . . . "

A. Capra pyrenaica pyrenaica .

B. ,, ,, lusitanica .

C. ,, ,, victoriae

D. ,, ,, hispanica .

IV. Capra ibex ...



V.



,, sibirica
A. Capra sibirica sibirica



B.

C.

D.

E.

F.

G.

H.

I.

J.

K.

L.

M.



fasciata
altaica
lydekkeri
hagenbecki
almasyi
merzbacheri
alaiana
wardi .
sakeen .
dauvergnei
pedri .
filippii .



VI. Capra nubiana . .

A. Capra nubiana nubiana

B. ,, ,, sinaitica

C. ,, ,, mengesi

VII. Capra walie .

VIII. hircus

A. Capra hircus segagrus

B. blythi .

C. ,, cretensis

D. picta

IX. Capra falconeri

A. Capra falconeri falconeri



B.
C.
D.
E.
F.



cashmiriensis

subsp.

megaceros .

jerdoni

chialtanensis, subsp. n.



Vol. I.
PAGK

131
131
133

134
136
137
138

138
139
139
140
140

141

142
143
144
144
145
145
146
147
148
148
149
150
151
152

153
154
154
155

155

156
157
159
161
161

161
163
164
166
167
169
171



V. Genus HEMITEAGUS.

I. Hemitragus jemlahicus

II. jayakari

III. ,, hylocrius



173
176
177



C.
1).
E.
F.
G.
H.
I.



GENERAL SYSTEMATIC INDKX x j x

Subfamily iii. RUPICAPRIN.E.

I. Genus RUPICAPRA. V PAGK'
Rupicapra rupicapra ..... 180

A. Rupicapra rupicapra rupicapra .... 181

B. .. ., faesula .... 182

C. .. .. ornata .... 182

pyrenaica . . . .183
parva . . . .183

F. .. caucasica . . . .183

G. .. ., asiatica .... 185

II. Genus CAPRICORNIS.

1. Subgenus CAPRICORNIS.
I. Capricornis sumatrensis ...... 188

A. Capricornis sumatrensis sumatrensis . . . 189

B. ,, . robinsoni 189



swettenhami . . . 189

milne-edwardsi . . 190

jamrachi . . .191

rubidus . . . .192

thar . . . .193

humei . 195



rodoni .... 195

II. Capricornis argyrochaetes ...... 196

2. Subgenus CAPRICORXULUS.

III. Capricornis (Capricornulus) crispus .... 200

A. Capricornis crispus crispus ..... 200

B. ,, ,, pryerianus .... 201

IV. Capricornis (Capricornulus) swinhoei . . . 201

III. Genus NEMORH.EDUS.

I. Nemorhoedus goral ....... 203

II. ., hodgsoni ... . 206

III. ., griseus ...... 206

IV. ., caudatus ...... 208

V. ., raddeanus 209

IV. Genus BUD!ORCAS.

I. Budorcas taxicolor , . . . . . .211

A. Budorcas taxicolor taxicolor . . . .211

B. whitei . ... 213

II. Budorcas tibetana ....... 214

III. bedfordi 217

V. Genus OREAMNOS.

Oreamnos americanus . . . . . . .218

A. Oreamnos americanus americanus . . . 219

B. ,, Columbia . . . 220

C. ,, ,, missoulae .... 220

D. ,, ,, kennedyi .... 221



XX GENERAL SYSTEMATIC INDEX

Subfamily iv.



Genus OVIBOS. P ^Q E ' PAG ' E



Ovibos moschatus ... . 224

A. Ovibos moschatus moschatus

B. wardi . . 227

C. ,, ,, niphcecus .... 228

D. ,, melvillensis .

E. ,, ,, mackenzianus . . . 229

Subfamily v. BUBALIN.E.

I. Genus B u B A L I s. ^'PA^E

I. Bubalis buselaphus . . 4

II. ,, major . . 6

A. Bubalis major major . ...

B. ,, ,, invadens . ... 100

C. matschiei . .. 10

III. Bubalis tora ...... . . 8

A. Bubalis tora tora . 10

B. digglei .

C. ,, ,, rahatensis . . .

D. swaynei

E. ,, ,, noacki .

IV. Bubalis cokei .....

A. Bubalis cokei cokei . . ... * .15

B. rothschildi .

C. ,, ,, nakurse ....

D. ,, ,, kongoni . . ... .17

Other names . . . . . . . 101

V. Bubalis neumanni . .18

VI. lelwel . . ... . 19 101

A. Bubalis lelwel lelwel ... .20

B. ,, , tschadensis . . . . . 20

B 1 . , modestus . . . ... 102

C. ,, , jacksoni . . . . .21

C 1 . , kenise . . ... 102

D. ,, , niediecki . . . . .23

E. , insignis ..... 23

F. , roosevelti . ... . 24

VP. Bubalis luzarchei . . . . , . . ..102

VII. caama . . . . . .24

A. Bubalis caama caama . . . .. . 27

B. selbornei . 27

VIII. Bubalis lichtensteini . . . .-" . . 29

II. Genus DAMALISCUS.

I. Damaliscus pygargus . . . . . . 33

II. albifrons .^ . . . . . 35

III. lunatus . 37



GENERAL .SYSTEMATIC INDEX



XXI



IV. Damiliscus korrigum

A. Dauialiscus korrigum korrigum .
A 1 . ,, lyra.

A 2 . .. 'purpurescens

B. ., tiang

C. ., selousi

D. .. jonesi
D 1 . .. topi .

E. ., jimela
E 1 .. eurus

F. .. phalius

V. Damaliscus hunteri .



Vol. II. V.
PA<;K PAOK

39 103

. 41

.. 103
. . 104

. 42

. 44

. 44

.. 104
44 104
.. 105

. 46

46



III. Genus CONNOCH.ETES.

I. Connochaetes gnu .....

II. ,, taurinus ....

A. Connochaetes taurinus taurinus .

B. , , johnstoni .
B l . , . cooksoni .

C. , , hecki

D. , , albojubatus

E. , , mearnsi .
Other names



50

54
56

57

57
58



105



106
106



Subfamily vi. CEPHALOPHIN.E.
Genus CEPHALOPHUS.
1. Subgenus CEPHALOPHUS.

I. Cephalophus sylvicultrix .

II. ,, jentinki .

III. ,, spadix ......

IV. ,, natalensis .

A. Cephalophus natalensis natalensis

B.

C.

D.

V. Cephalophus centralis

VI. ,, nigrifrons

VII. claudi .

VIII. rubidus

IX. ,, weynsi

X. ,, johnstoni

XI. ignifer

XII. harveyi



amcenus
robertsi .
bradshawi



A. Cephalophus harveyi harveyi

B. ,, ,, keniae

XIII. Cephalophus leopoldi .

XIV. ,, leucogaster



63
66
67

68
69
69
70
71

71
72
73
73
74
75
75

76

77
77

78
79



XXII



GENERAL SYSTEMATIC INDEX



Vol. II.



XV. Cephalophus dorsalis .

XVI. ,, castaneus .

A. Cephalophus castaneus castaneus

B. ,, ,, orientalis

XVII. Cephalophus leucochilus

(?) ,, emini . .

XVIII. ogilbyi .

XIX. brookei.

XX. ,, callipygus

XXI. ,, rufilatus

A. Cephalophus rufilatus rufilatus .

B. ,, ,, rubidior

XXII. Cephalophus doriae . . .

XXIII. niger .

XXIV. walked .



PAGE PAGE
80

81

.. 106

.. 106

82
83
83
84
85

86
87



91
92



2. Subgenus GUEVEI.

XXV. Cephalophus (Guevei) maxwell! .

XXVI. ,, ,, melanorheus

A. Cephalophus melanorheus melanorheus

B. ,, sundevalli .

C. ,, anchietss

D. sequatorialis
D 1 . ,, schultzei .

E. musculoides

XXVII. Cephalophus (Guevei) monticola

XVIII. nyasai .

A. Cephalophus nyasas nyasae . . .

B. ,, ,, defriesi

C. ,, ,, congicus

XXIX. Cephalophus (Guevei) hecki . ;

XXX. ,, ,, simpsoni .

XXXI. ,, ,, lugens

3. Subgenus SYLVICAPRA.

XXXII. Cephalophus (Sylvicapra) grimmi
A. Cephalophus grimmi grimmi

B ,, flavescens . .

C. altifrons . ;

D. splendidulus

E. ,, coronatus .

F. , campbelliee
G-, , abyssinicus .
G 1 . , pallidior

G 2 . , luteus . ....

H. , nyansae

I. , hindei

J. 5 altivallis



107



93

96
97
97



99
100

103
103
104
104

105
105

106



107

110

112

113

114

114

116

117 107
.. 107
.. 108

118

119

120



C.


'


nai uaui IA.UIU

aureus


D.


j


schillings!


E.


j


somalicus .


F.


.


aceratos


G.


,


porteousi .



<;KNKI;AI. SYSTEMATIC INDEX xxui

Vol. II. V.
PAGE PAGE

K. Cephalophus grimmi shirensis .... 120
L. .. ., roosevelti .... 121


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