Plantae Wilsonianae; an enumeration of the woody plants collected in western China for the Arnold arboretum of Harvard university during the years 1907, 1908, and 1910 (Volume 1:1) online

and soon separating into thin flakes. Wilson's Nos. 4391 and 4899 seem nearest
to the type. No. 4484 is also near the type, except that it has the leaves loosely
pubescent beneath, while the other numbers of Wilson differ in their large foliage,



SAXIFRAGACEAE. HYDRANGEA 27

the leaves attaining a length of 15-20 cm. with the petioles 4 cm. long and the
inflorescence measuring 25 cm. hi diameter.

Hydrangea xanthoneura, var. Wilsonii Rehder, n. var.

A typo recedit ramulis hornotinis griseo-flavescentibus, annotinis
griseis v. avellaneis lenticellis pallidis instructis cortice adhaerente et
foliis junioribus subtus sparse adpresse pubescentibus. Arbor ad 5 m.
alta.

Western Szech'uan: Ta-p'ao-shan, northeast of Tachien-lu,
woodlands, alt. 2400-2800 m., July 3, 1908 (No. 2407, type); Pan-
lan-shan, west of Kuan Hsien, thickets, alt. 2200 m., June 26, 1908
(No. 2410); Wa-ssu country, W6n-chuan Hsien, thickets, alt. 1600-
2400 m., July 1908 (No. 2408).

The color and behavior of the bark would seem to afford good specific differ-
ences, if it were not for the following specimens which are intermediate in this re-
spect between the type and the variety, but are apparently closer to the variety.

Hydrangea xanthoneura, var. glabrescens Rehder, n. comb.

Hydrangea serrata Koehne, Deutsch. Dendr. 189 (not De Candolle) (1893).
Hydrangea Bretschneideri, var. glabrescens Rehder hi Bailey, Cycl. Am. Hort.
II. 784 (1900); in Mitt. Deutsch. Dendr. Ges. XII. 121 (1903).

A typo recedit foliis minoribus, tenuioribus et plerumque angusti-
oribus, saepe grossius serratis, venis supra non flavidis, subtus fere
glabris v. sparse pubescentibus nee strigosis.

Seed collected by Dr. E. Bretschneider near Pekin and introduced
into cultivation: Hort. Bot. Berol.; E. Koehne, Herb. Dendr. (No.
113); Arnold Arboretum, in Herb. Arnold Arboretum.

It is with some hesitation that I refer this variety to H. xanthoneura, but M the
pubescence is to be considered the chief distinguishing character between this
species and H. Bretschneideri, it must be referred to the former.

Western Szech'uan: Mupin, thickets, alt. 1500-2500 m., June and
October 1908 (Nos. 1183, 1327); thickets around Tachien-lu, alt.
2100-2400 m., October 1908 (No. 1347) ; near Tachien-lu, 2700-3600 m.,
A. E. Pratt (No. 285). Western Hupeh: Hsing-shan Hsien,
thickets, alt. 1500-1800 m., July 1907 (No. 2398); no locality, June
1901 (Veitch Exped. No. 2184). Yunnan: Mengtze, alt. 2700 m.,
A. Henry (No. 10235).

Hydrangea pubinervis Rehder, n. sp.

Frutex ad 3 m. altus ramulis junioribus glabrescentibus purpureis,
vetustioribus castaneis lenticellis sparsis instructis. Folia mem-
branacea, alte elliptica, acuminata, basi late cuneata, grossius ser-



28 WILSON EXPEDITION TO CHINA

rato-dentata, supra flavo-viridia, glabra v. fere glabra nervis strigosis
exceptis, subtus pallidiora, tota facie crispulo-villosa ad nervos pilis
flavidis adpressis dense obtecta, 6-9 cm. longa et 3.5-4.5 cm. lata;
petioli graciles, 1.5-2 cm. longi, pubescentes. Cyma fere plana, e
radiis 5-7 decussatis distantibus composita, strigosa; flores radiantes
circa 2.5 cm. diam. sepalis 4 late ovalibus v. obovatis; calycis denies
riorum fertilium triangulares; petala oblongo-ovata, apice cucullata,
2 mm. longa; stamina inaequalia, minora petalis breviora, majora
petalis paullo longiora; ovarium semisuperum; stylis 3 erectis. Cap-
sulae maturae desiderantur.

Western Szech'uan: Wa-shan, thickets, alt. 1500-2250 m.,
July 1908 (No. 2411).

Closely related to H. xanthoneura Diels which is easily distinguished by the leaves
being serrulate, narrower and larger and on the under side nearly glabrous except
the strigose veins.

Hydrangea Bretschneideri Dippel, var. setchuenensis Rehder, n.
var.

A typo recedit foliis majoribus ad 20 cm. longis et 11 cm. latis, venis
saepe flavescentibus, petiolis densius pubescentibus, ramulis juniori-
bus laxe et sparse pubescentibus, annotinis pallide brunneis lenticellis
conspicuis instructis, cymis usque ad 25 cm. diam., laxis, convexis,
interdum fere paniculiformibus.

Western Szech'uan: Wa-ssu country, Wen-chuan Hsien, thickets,
alt. 2300-2700 m., July and November 1908 (No. 1323). Western
Hup eh: Fang Hsien, rocky places, alt. 1800 m., August 1907 (No.
2399).

The Szech'uan and the Hupeh plants differ slightly; the former has the leaves
generally ovate-oblong and approaches also in the somewhat darker color of the
branchlets the type, while the leaves of the Hupeh plant are elliptic-oblong or
oblong-obovate.

Hydrangea Bretschneideri, var. lancifolia Rehder, n. var.

A typo recedit ramulis cinereis lenticellis conspicuis instructis,
foliis oblongo-lanceolatis, supra sparse strigosis, subtus tota facie
dense crispulo-villosis et ad nervos strigosis, 7-12 cm. longis et 2-3
cm. latis.

Western Szech'uan: Tachien-lu, thickets, alt. 2100-2400 m.,
June 1908 (No. 2412).

This variety might possibly be considered a pubescent and narrow-leaved form
of H. xanthoneura, since it has almost the same kind of bark as var. Wilsonii of
that species.



SAXIFRAGACEAE. HYDRANGEA 29

Hydrangea Sargentiana Rehder, n. sp.

Frutex robustus bimetralis; ramuli juniores dense villosi et excre-
scentiis crebris e trichomatibus basi confluentibus purpureis formatis
praediti. Folia ovato-oblonga, acuminata, basi rotundata v. interdum
subcordata v. late cuneata, inaequaliter et fere dupliciter ciliato-
dentata dentibus setaceoacuminatis, 15-30 cm. longa et 6.5-16 cm.
lata, supra obscure viridia, scabro-strigosa, subtus dense villoso-hirta,
ad nervos hirto-pilosa excrescentiis purpureis intermixtis; petioli
dense pilosi et excrescentiis purpureis instructi, 3-9 cm. longi. Cyma
fere plana, 12-16 cm. diam., densa, dense pubescens, e radiis oppo-
sitis congestis 7-13 composita; flores radiantes graciliter pedicellati
2.5-3.5 cm. diam. albi, sepalis 4, orbiculato-obovatis integris; sepala
florum fertilium late triangularia, tubo dimidio breviora; petala ovata,
mox caduca, apice interdum calyptratim coherentia, 2 mm. longa;
stamina breviora petalis aequilonga, longiora circa 4 mm. longa;
styli plerumque 3 v. interdum 2. Capsula hemispherica, leviter 10-
12-costata, apice truncata, 3 mm. diam.; semina elliptica, utrinque in
alam brevem contracta, striata, flavo-brunnea, circa 0.7 mm. longa.

Western Hupeh: Hsing-shan Hsien, thickets, alt. 1500-1800 m.,
August and December 1907 (No. 772).

Adistinct species with large and handsome foliage similar to H. robusta Hooker
f . & Thomson, which is easily distinguished by its strigose pubescence, ovate
leaves, by the serrate sepals of the sterile flowers and the larger capsule. The allied
H. Rosthornii Diels also differs in its strigose pubescence and ovate leaves.

Hydrangea villosa Rehder, n. sp.

Frutex 1-3 m. altus ramulis junioribus albido- v. fulvo-villosis
angulatis. Folia elliptico- v. oblongo-lanceolata, acuminata, basi
cuneata, fimbriato-denticulata dentibus setaceo-acuminatis, 10-20
cm. longa et 3.5-6.5 cm. lata, supra obscure flavo-viridia, scabro-
strigosa pilis basi bulbosis, ad costam mediam pilis longioribus al-
bidis instructa, subtus pilis patentibus hirto-villosis incano-tomentosa,
ad nervos pilis ad 3 mm. longis et saepe, praesertim ad costam mediam,
fulvescentibus v. fulvis praedita; petioli villosi saepe pilis fulvis longis
patentibus interspersis, 1^ cm. longi. Cyma convexa, interdum
plana, albido- v. fulvo-villosa, radiis oppositis 7-9 satis remotis
composita; flores radiantes graciliter pedicellati, 3^ cm. diam., coeru-
lescentes, sepalis 4 orbiculato-obovatis plerumque crenulatis; pedi-
celli florum fertilium strigosi, tubum calycis glabri v. basi tantum
strigosi subaequantes, dentibus triangularibus vix dimidium tubum
aequantibus; petala oblongo-ovata, acutiuscula, concava, patentia,



30 WILSON EXPEDITION TO CHINA

2 mm. longa; stamina valde inaequalia, minora petalis subaequi-
longa, majora eis duplo longiora; styli 2, patentes, apice incrassati.
Capsula hemispherica, apice truncata, leviter costata, 2.5-3 mm.
diam.; semina late elliptica, utrinque in alam constricta, striata,
flavido-brunnea, circa 0.6 mm. longa.

Western Szech'uan: Wa-ssu country, Wn-chuan Hsien, thick-
ets, alt. 1800 m., July and November 1908 (No. 1227, type), alt. 1200-
2700 m., July and November 1908 (No. 1250), October 1910 (No.
4302); Pan-lan-shan, west of Kuan Hsien 1 , woodlands, alt. 2400 m.,
August 1910 (No. 4483).

Easily distinguished from allied species by its villous pubescence.

Hydrangea glabripes Rehder, n. sp.

Frutex metralis ramulis glabris v. fere glabris, vetustioribus flavo-
griseis. Folia lanceolata, acuminata, basi late cuneata v. rotundata,
dentato-serrulata serraturis mucronulato-acuminatis, 6-12 cm. longa
et 2-3 cm. lata, supra obscure flavo-viridia, satis dense strigulosa,
subtus pallida, hirto-villosa costa fere glabra excepta; petioli graciles,
1.5-4 cm. longi, glabri, tantum supra in canaliculo parce pubes-
centes. Cyma fere plana v. leviter convexa, 8-10 cm. diam., strigosa,
radiis brevibus 7-9; flores radiantes pauci, rosei, sepalis 4, late obo-
vatis, leviter emarginatis integris; flores fertiles rosei; calycis tubus
hemisphericus, glaber ima basi excepta, dentibus triangularibus min-
utis; petala oblongo-ovata, 1.5 mm. longa; stamina inaequalia, longi-
ora 4 mm. longa; styli 2. Capsulae maturae desunt.

Western Hupeh: Fang Hsien, thickets, alt. 1200-1800 m., August

1907 (No. 2391).

Closely related to H. longipes Franchet which is chiefly distinguished by broader
strigose or glabrescent leaves, strigose branchlets, longer petioles and white flowers.
From all allied species H. glabripes differs in the leaves having a densely villous
under side and at the same time a glabrous petiole and a glabrous or nearly gla-
brous midrib.

Hydrangea aspera D. Don, var. velutina Rehder, n. var.

Differt a typo foliis minoribus subtus densius velutino-hirtellis,
floribus fertilibus sterilibusque coeruleis v. roseis minoribus, stylis
plerumque 2. Folia oblongo-ovata v. ovato-lanceolata, fimbriato-
denticulata, 8-12 cm. longa et 3-5 cm. lata; petioli 1-2 cm. longi;
petala, stamina, styli coerulea; flores radiantes steriles 3 cm. diam.,
sepalis crenato-serratis.

Western Szech'uan: Wa-shan, thickets, alt. 1800-2400 m., July

1908 (Nos. 2405, 2403 (floribus omnibus sterilibus)); Mupin, thickets,



SAXIFRAGACEAE. HYDRANGEA 31

alt. 1200-1500 m., June 1908 (No. 2404); Lungan Fu, thickets, alt.
1800-2700 m., August 1910 (No. 4482).

This form is probably best treated as a variety of H. aspera, though it shows a
close relation to H. villosa; it lacks, however the villous and fulvous hairs of that
species. In No. 2404 and 4482 the pubescence is less velutinous and approaches
that of H. strigosa.

Hydrangea aspera, var. scabra Rehder, n. var.

A typo recedit praecipue foliis ovatis v. ovato-oblongis, basi rotun-
data v. subcordata, supra manifeste scabra, subtus dense velutino-
hirtellis, 12-18 cm. longis et 7.5-10 cm. latis. Capsula circa 3 mm.
diam., stylis 2.

Western Szech'uan: southeast of Tachien-lu, thickets, alt.
2400 m., October 1910 (No. 4485).

In the shape of the leaves this variety approaches somewhat H. Rosthornii Diels,
but is easily distinguished by the nearly velutinous pubescence of the under side
of the leaves and by the shorter petioles.

Hydrangea strigosa Rehder, n. sp.

H. aspera Hemsley in Jour. Linn Soc. XXIII. 272 (not D. Don) (1887).

Pampanini in Nuov. Giorn. Bot. lial n. ser. XVII. 283 (1910).
H. aspera, f. typica Diels hi Bot. Jahrb. XXIX. 375 (1900).

Frutex 2-3 m. altus, ramulis junioribus teretibus v. leviter angu-
latis strigosis, vetustioribus griseo-brunneis lenticellis destitutis.
Folia oblonga v. oblongo-lanceolata, acuminata, basi cuneata, denti-
culato-serrata serraturis porrectis parvis mucronulatis, 8-23 cm.
longa et 2-6 cm. lata, supra sparse strigosa, subtus densius strigosa,
praecipue ad venas; petioli 1.5-3 cm. longi, dense strigosi. Cyma
convexa, radiis 5-7, dense strigosa; flores radiantes graciliter pedi-
cellati, sepalis late ovalibus plerumque mucronulatis integris v.
sparse serrulatis; flores fertiles albi; sepala triangularia, calycis di-
midium tubum parce strigosum vix aequantia, glabra v. fere glabra;
petala ovato-oblonga, 2 mm. longa; stamina longiora circa 6 mm.
longa, breviora dimidio breviora; styli 2. Capsula hemispherica,
truncata, 3 mm. alta, costata; semina late elliptica, utrinque subito
in alam contracta, striata, 0.6 mm. longa, flavo-brunnea.

Western Hupeh: north and south of Ichang, thickets, alt. 500-
1200 m., August and December 1907 (No. 765, type): Hsing-shan
Hsien, thickets, alt. 900-1500 m., August and December 1907 (Nos.
773, 2394); Patung Hsien, thickets, alt. 1200-1500 m., August 1907
(Nos. 2392, 2395); Fang Hsien, thickets, alt. 1200-1800 m., August
1907 (No. 2390, with all the flowers sterile); Packang, 1901 (Veitch



32 WILSON EXPEDITION TO CHINA

Exped. No. 1473); South Wushan, 1901 (Veitch Exped. No. 2446);
without locality, 1901 (Veitch Exped. No. 2527); A. Henry (No.
1083). Western Szech'uan: Mt. Omei, 1904 (Veitch Exped. No.
4902 with all the flowers sterile); Nanch'uan, Shan-tzu-p'ing, A. von
Rosthorn (No. 299).

Closely related to H. aspera D. Don which chiefly differs in the fimbriate-denticu-
late leaves being on the under side on the veins and veinlets covered with a villose-
hirtellous, not appressed strigose pubescence and in the larger capsules with
usually three styles. The difference in pubescence and serration, serrulate, with the
teeth pointing forward in //. strigosa, and denticulate or dentate with spreading
teeth in H. aspera, seems to distinguish clearly the Chinese from the Himalayan
plant of which I have seen Wallich's No. 440 B; the pubescence of that specimen
agrees well with Don's original description "foliis . . . subtus dense cano-tomen-
tosis " and Clarke's description "under surface with gray woolly hairs," terms
which hardly could be applied to the strigose pubescence of the Chinese plant. In
size and shape of foliage the Chinese plant seems exceedingly variable, and the fol-
lowing forms pass gradually into each other.

Hydrangea strigosa, var. macrophylla Rehder, n. comb.

H. aspera, var. macrophylla Hemsley in Jour. Linn. Soc. XXIII. 273 (1887).
Diels in Bot. Jahrb. XXIX. 375 (1900).

Western Hupeh: A. Henry (Nos. 2083, 6477); Fang Hsien, thick-
ets, alt. 1300 m., August 1907 (No. 571); Hsing-shan Hsien, thicket,
alt. 900-1200 m., August and December 1907 (No. 757). Western
Szech'uan: Mt. Omei, July 1904 (Veitch Exped. No. 4902); Nan-
ch'uan, Chang-kou, September 18, 1891, A. von Rosthorn (No. 953).

The leaves of this form are sometimes 28 cm. in length and 11 cm. in width and are
generally ovate-lanceolate or sometimes ovate-oblong and nearly rounded at the
base. This form passes gradually into the type, and some specimens enumerated
under the type, as Wilson's Nos. 2394 and also No. 773 and Henry's No. 1083,
with leaves 22 cm. long and 6.5 cm. broad, might perhaps be referred to var.
macrophylla, but the leaves are comparatively narrow and cuneate at the base.

Etydrangea strigosa, var. sinica Rehder, n. comb.
H. aspera, var. * sinica Diels in Bot. Jahrb. XXIX. 375 (1900).

Western Hupeh: A. Henry (No. 185); Nan-t'o, A. Henry (No.
2206) . WesternSzec h'u a n : Nanch'uan, Ma-f ou-lin-p'o, August 26,
1901, A. von Rosthorn (No. 629).

Differs in its oblanceolate or oblong-obovate leaves rarely exceeding 10 cm.
in length, the leaves having generally their greatest width above the middle, not
below as in the other forms.

Hydrangea strigosa, var. angustifolia Rehder, n. comb.
H. aspera var. angustifolia Diels 1. c.



SAXIFRAGACEAE. - 1 - HYDRANGEA 33

Western Szech'uan: Wang-t'ien-ling, August 9, 1891, A. von
Rosthorn (No. 1168, type). Hupeh: Changyang Hsien, thickets,
alt. 30-60 m., September 1907 (Nos. 2393, 2396).

Wilson's No. 2396 with the leaves 23 cm. long and 3.7 cm. broad approaches
var. macrophylla, but on account of the narrowness of the leaves it finds its place
perhaps better here.

Hydrangea Rosthornii Diels in Bot. Jahrb. XXIX. 374 (1900).

Western Szech'uan: Nanchuan, Ya-chih-pa, August 1891, A. von
Rosthorn (No. 471, type); Wa-shan, thickets, alt. 1500-2400 m., July,
September and November 1908 (Nos. 1156, 1348, 1372, 2414); Ta-
cbien-Iu, thickets, alt. 1800-2500 m., July and October 1908 (No.
1365), October 1910 (No. 4337); Mupin, thickets, alt. 2400 m., October
1910 (No. 4341); Mt. Omei, July 1904 (Veitch Exped. No. 4903).

The close relationship of this species with H. robusta Hooker f . & Thomson is
still more evident in Wilson's specimens than in the type specimen; the leaves of
Wilson's specimens are larger, the largest measuring 25 cm. in length and 17 cm. in
width, the longest petiole is 20 cm. long and in some specimens the sterile flowers
have strongly toothed sepals. It is also closely related to H. longipes Franchet,
which is chiefly distinguished by its smaller and thinner, coarsely serrate leaves,
less densely strigose or glabrescent beneath.

Hydrangea longipes Franchet in Nouv. Arch. Mus. Paris, se*r. 2,
VIII. 228 (PL David. II. 45) (1885).

Hydrangea ( Calyptranthe) longipes Hemsley in Jour. Linn. Soc. XXIII. 273

(1887).

Hydrangea aspera, var. a longipes Diels in Bot. Jahrb. XXIX. 374 (1900).
Hydrangea (Calyptranthe) Hemsleyana Diels L c. 376 (1900). Pampanini in

Nuov. Giorn. Bot. Ital. n. ser. XVII. 284 (1910).

Western Szech'uan: Mupin, June 1869, P. David] Mupin, thick-
ets, alt. 1800 m., July 1908 (No. 2406); alt. 1500-2100 m., October
1910 (No. 4342); Wa-ssu country, Wen-chuan Hsien, thickets, alt.
1100-2100 m., July 1908 (No. 2400); Hsing-shan Hsien, thickets, alt.,
1200-1800 m., July 1808 (No. 2401); Wa-shan, thickets, 1800-2400
m., July 1908 (No. 2413); northeast of Sungpan, thickets, alt. 1800-
2200 m., October 1910 (No. 4330); Lungan Fu, alt. 1800-2400 m.,
August 1910 (No. 4481); Nanch'uan, A. von Rosthorn (No. 1931).
Western Hupeh: Chang-lo Hsien, cliffs and thickets, alt. 1300-
1500 m., July 1907 (No. 2402); north and south of Ichang, thickets
and rocky places, alt. 1200-1800 m., June and November 1907 (No.
580); Patung, May 1901 (Veitch Exped. No. 1213); without locality,
August 1901 (Veitch Exped. No. 2514); A. Henry (Nos. 5839 A,
5839 B).



34 WILSON EXPEDITION TO CHINA

Hydrangea longipes is closely related to H. Giraldii Diels and also to H. robusta
Hooker f . & Thomson as Franchet has already stated, and some of the specimens,
as Wilson's No. 580, which has rather densely pubescent and large leaves, seem
somewhat intermediate. Hydrangea longipes Franchet and H. longipes Hemsley are
identical; the apparent discrepancies in the descriptions do not exist in reality.
First it is to be stated that H. longipes Hemsley does not belong in the section
Calyptranthe where Hemsley placed it, apparently misled by the fact that the
petals occasionally cohere at the apex, but this may happen in almost all species
of the subsection Asperae, which does not differ in the structure of its flowers from
Calyptranthe. The description of Franchet contains some misleading inaccuracies;
the flowers have 10 stamens and 2 styles, not 5 stamens and 3 styles as Franchet
describes them, and the leaves of at least part of the specimens collected by David
attain a length of four inches, while Franchet says "folia vix ultra-pollicaria."
The only difference I can find between Franchet's and Hemsley's specimens is the
somewhat slighter pubescence of the latter, the leaves being sometimes nearly
glabrous. A form based on No. 846 of C. Silvestri from the mountains of Chia-
yuen-kou, western Hupeh, with the radiant flowers smaller and greenish has been
described by Pampanini as H. Hemsleyana, var. Pavoliniana (Nuov. Giorn. Bot.
Ital. n. ser. XVII. 284 [1910]).

Hydrangea anomala D. Don, Prodr. Fl. Nepal 211 (1825).

Hydrangea altissima Wallich, Tent. Fl. Nepal t. 50 (1826).
Hydrangea glabra, Hayata in Jour. Coll. Sd. Tokyo, XXV. 89, t. 6 (FL Mont.
Formos.) (1908).

Western Hupeh: north and south of Ichang, common, alt. 1200-
1800 m., May and October 1907 (No. 491, in part); Packang, July
1901 (Veitch Exped. No. 894); A. Henry (No. 6511). Western
Szech'uan: Wa-shan, woodlands, alt. 1800-2500 m., June and Octo-
ber 1908 (No. 491, in part); Mt. Omei, July 1904 (Veitch Exped. No.
4901); Nanch'uan, Yang-yii-ping, A. von Rosthorn (No. 51); South
Wushan, A. Henry (Nos. 5557, 5658). Formosa: Mt. Morrison, T.
Kawakami & U. Mori (No. 1723 ex Hayata).

I have been unable to detect in Hayata's description and plate any character
by which to distinguish his H. glabra from H. anomala.

SYNOPSIS OF THE CHINESE SPECIES OF HYDRANGEA.

As the determination of the copious material collected by Wilson of this genus
has necessitated a study of all the Hydrangeas of China and the adjacent parts of
Asia, it may be useful to place the results of this study permanently on record in
the shape of a synopsis of all the Chinese species of Hydrangea.

KEY TO THE SPECIES.

* Semina exalata v. utrinque in alam protracta; petala apice soluta, patentia;
frutices erecti raro scandentes ramulis annotinis medulla alba solida ampla.

Sect. I. EUHYDRANGEA.

t Ovarium semisuperum; capsula ovoidea apice in stylos plerumque 3 attenu-
ata; stamina petala vix v. paullo superantia, subaequalia.



SAXIFEAGACEAE. HYDRANGEA 35

J Semina exalata v. alls brevissimis instructa; cyma plana, rarius convexa;
petala plerumque tarde decidua, sub anthesi reflexa.

Subsect. 1. PETALANTHAE.
Cyma basi foliis suffulta; pars libera capsulae tubo calycis longior; folia

denticulata v. denticulata-serrata.

Cyma umbellam sessilem formans (rarius ramulis oppositis remotis
simulque foliis glabris); capsula ovoidea longior quam lata; petala
obovata.
Folia glabra, subtus viridia, chartacea v. coriacea; flores radiantes

pauci v. deficientes 1. H. chinensis.

Folia subtus glaucescentia sparse ad venas densius pubescentia;

cyma manifeste umbelliformis 2. H. umbellate.

Cyma e ramulis oppositis plus minus remotis composita; folia ad venas
saltern pubescentia; flores fertiles coerulei; petala oblongo-obovata
v. lanceolata.

Sepala anguste lanceolata; petala lanceolata; styli recti v. leviter
apice curvati; capsula subglobosa, absque styli latior quam

longa 3. H. Davidii.

Sepala triangulari-ovata; petala obovato-oblonga; styli apice revo-
luti; folia plerumque oblanceolatft-oblonga; capsula ovoidea, absque

stylis paullo longior quam lata . ' 4. H. yunnanensis.

Cyma pedunculata aphylla; pars libera capsulae tubo calycis brevior,

rarius aequilonga; folia argute serrata; flores fertiles coerulei v. rosei.

Folia utrinque et ramuli strigillosa pilis basi bulbosis; flores radiantes

disepali pauci 5. H. Moellendorfii.

Folia utrinque glabra v. supra sparse pilosa et subtus secus venas vil-
losa; ramuli glabri; flores radiantes quadrisepali.

6. H. opuloides.

tt Semina utrinque in alam protracta; flores fertiles albi; cyma corymbiformis
et convexa v. paniculiformis; petala cito decidua.

Subsect. 2. HETEROMALLAE.

Inflorescentia manifeste paniculata; folia subtus ad venas tantum pu-
bescentia v. glabrescentia 7. H. paniculata.

Infloreseentia cymosa plerumque convexa.

Folia subtus glabrescentia, ad venas tantum v. interdum ad venulas

sparse pubescentia.
Folia subtus glauca, sub lente dense papillosa, ad venas pilis sericeis

dense obtecta, basi rotundata 8. H. hypoglauca.

Folia subtus pallide viridia sine papillis, ad venas strigosa, basi plus

minus attenuata 9. H. xanthoneura.

Folia subtus tota facie pubescentia.
Folia subtus pubescentia tantum.

Folia subtus ad costam mediam venasque dense pilis flavidis ad-
pressis obtecta, ceterum laxe villoso-strigosa, elliptica, grossius

serrata 10. H. pubinervis.

Folia subtus cinereo-villosa, plerumque ovato-oblonga, denticu-

lato-serrulata 11. H. Bretschneideri.

Folia subtus dense albo-tomentosa, denticulato-serrulata.

12. H. mandarinorum.
ft Ovarium totum inferum; capsula hemispherica v. turbinata stylis plerumque

duobus Subsect. 3. ASPERAE.

t Ramuli petiolique et saepe costa subtus pilis longis patentibus instructa v.
tomentosa.



36 WILSON EXPEDITION TO CHINA

Pubes ramulorum petiolorum costarum subtus trichomatibus carnosis,
aculeiformibus apice fissis plerumque purpureis crebris interspersa;
folia ad 25 cm. longa, basi plerumque rotundata v. subcordata.

13. H. Sargentiana.
Pubes tantum pilosa v. tomentosa.

Capsula et sepala utrinque tomentosula; folia oblongo-lanceolata, ad
20 cm. longa, supra scabra, subtus ad costas tomentosa.

14. H . longifolia.