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Samuel Hubbard Scudder.

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as long as broad.

g l . Hind tibiae of $ at least a tenth longer than the
femora.

h l . Body without conspicuously contrasted colors ;
hind femora of relatively slender, four times as
long as broad 36. occultus.



SCUDDER. NORTH AMERICAN CEUTHOPHILI. 29

h 2 . Body with conspicuously contrasted colors; hind
femora of <$ relatively slender, less than three and
a quarter times as long as broad . 46. discolor.

g 2 . Hind tibiae of $ less than one tenth longer than

the femora.

A 1 . Hind femora of $ with no raised points on the
upper distal half.

1 1 . Outer carina of hind femora of $ almost
unarmed ; markings of the body more or less
marmorate or maculate.

J 1 . A broad continuous light dorsal stripe on
pronotum, usually extending over the whole
thorax.

k 1 . Hind tibial spurs distinctly marked with
black at base ; ovipositor twice as long as

fore femora 27. latens.

k 2 . Hind tibial spurs at most indistinctly
infuscated at base ; ovipositor shorter than

fore femora 31. tenebrarum.

j 2 . A narrow and very unequal light dorsal
stripe on pronotum, interrupted^ if present, on
rest of thorax 38. bruneri.

1 2 . Outer carina of hind femora of $ finely and
closely serrate ; dark markings of body confined
to transverse borderings of the segments.

48. vinculatus.

h 2 . Hind femora of $ with a greater or less num-
ber of raised points on upper distal half.
i 1 . Hind femora of $ with only a few distant
recumbent spines on outer carina.
j 1 . Hind tibiae of $ a tenth longer than the
femora; spurs fully twice as long as tibial

depth 44. pinguis.

j 2 . Hind tibiae of $ less than a tenth longer

than the femora ; spurs much less than twice

as long as tibial depth . . 40. mexicanus.

i?. Hind femora of $ with numerous denticula-

tions on the outer carina, forming a more or less

close serration.

j 1 . Ovipositor relatively short, at most bu(
little more thau half as long as hind femora.



30 PROCEEDINGS OP THE AMERICAN ACADEMY.

k\ Hind femora of $ less than twice as
long as fore femora.

P: Hind tibiae of <J no longer than
femora ; spurs only a little longer than
tibial depth, and divaricating about 60 ;
inner carina of fore femora minutely ser-
ratulate .... 50. californianus.
I 2 . Hind tibiae of $ a little longer than
femora ; spurs fully twice as long as tibial
depth, and divaricating about 90 ; inner
cariua of fore femora simply spined.

49. testaceus.

k 2 . Hind femora of $ two and a fourth times
as long as fore femora . . 29. neglectus.
f. Ovipositor relatively long, two thirds as
long as hind femora or more.

k l . Hind femora of $ relatively slender,
at least three times as long as broad.

I 1 . Hind tibiae of $ of same length as
femora ; colors moderately dark.

37. alpinus.

P. Hind tibiae of $ considerably longer
than femora; colors rather pallid.

41. pattescens.

k*. Hind femora of $ relatively stout,
hardly more than two and a half times as

long as broad 43. crassus.

c 2 . Dorsal surface of abdomen of $ closely tuberculate ; hind
tibiae strongly arcuate.

d l . Both outer and inner carinae of hind femora of $ armed
with a large compressed spine as long as the depth of the

genicular lobes 52. pacijlcus.

c? 2 . Outer carina of hiud femora of $ elevated to a high
lamina, suddenly terminating acutely before the genicular

lobes * 53. henshawi.

a 2 . Second joint of hind tarsi but little longer than the third.

b\ Large species ; outer carina of hind femora considerably and

uniformly elevated throughout 54. devius.

b z . Small species ; outer carina of hind femora elevated distally

much more than proximally 55. neomexicanus.

The male being unknown to me, G, sylvestris does not appear in the
above table. It will be found below as No. 41.



SCUDDER. NORTH AMERICAN CEDTHOPHILI. 31

1. CKUTHOPHILUS VARIEGATUS, sp. nov.

Blackish fuscous with a slight olivaceous tinge, marked with yel-
lowish luteous ; there is an interrupted dorsal stripe of the lighter
color especially interrupted on the anterior half of the pronotum ; this
is heavily margined by subconfluent dark blotches or spots, and the
dark color prevails over the rest of the thorax, with oblique patches of
the lighter tints on the meso- and metanotutn, irregular vermiculate
blotches of greater or less extent on the pronotum and the lower
edges of the descending lobes of the thorax margined not very nar-
rowly with luteous ; the sides of the abdomen are prevailingly luteous,
but the anterior outer margins are mostly fuscous ; the hind femora
have the usual scalariform markings very pronounced, while the other
legs are prevailingly luteous and much streaked with fuliginous,
excepting the tarsi ; the longer spines are bright luteous, but black
tipped. The antennae are two or three times as long as the body}
moderately stout at the base and gradually tapering, and the legs
moderately short. Fore femora distinctly broader than the middle
femora, nearly half as long again (() or scarcely a fourth as long again
( 9 ) as the pronotum, and less than half as long as the hind femora,
the inner carina with three long spines, the distal subapical and very
long. Middle femora with 2-3 long spines on the front carina, one
subapical and very long, and on the hind carina 1-2 long spines besides
a long genicular spine. Hind femora slightly (c?) or considerably (?)
more than twice as long as the fore femora, very broad and stout, dis-
tinctly less than three times as long as broad especially in the male, a
few very distant and scattered raised points over the whole apical half of
the surface, excepting beneath and especially on the inner side above,
the outer carina with 3-4 very unequal spines in the apical half, one,
sometimes two, much larger than the rest, coarse and as long as the
tibial spurs ( $ ) or with a single inconspicuous spine in the pregenicu-
lar sinus (9), the inner carina with half a dozen small uniform but
irregularly distant spinules, the intervening sulcus rather narrow.
Hind tibiae straight in both sexes, moderately slender, scarcely ex-
panded distally, distinctly but not greatly longer than the femora,
armed beneath with a single subapical spine besides the apical pair ;
spurs subalternate, the basal before the end of the proximal fourth of
the tibia, half as long again as the tibial depth, set at an angle of 40-45
with the tibia and divaricating about 50-60, their tips considerably
incurved ; inner middle calcaria a little larger than the outer, twice as
long as the others or as the spurs, and nearly as long as the first tarsal



32 PROCEEDINGS OP THE AMERICAN ACADEMY.

joint. Hind tarsi considerably less than half as long as the tibia, the
first joint unusually prolonged below and as long as the rest together,
the second fully twice as long as the third and with it as long as the
fourth. Cerci moderately stout, regularly tapering, as long as the
pronotum. Ovipositor very short, not so long as the pronotum, rap-
idly tapering at base, beyond slender, the armature of the inner valves
consisting of moderately stout but rather prominent bluntly pointed
spines.

Length of body, $ 15 mm., 9 19 mm.; antennas (est.), $ 30+
mm., 9 45+ mm.; prouotum, $ 5.5 mm., 9 6.5 mm.; fore femora,
$ 8 mm., 9 7.75 mm.; hind femora, $ 16.75 mm., 9 17.75 mm. ;
hind tibia?, $ 19 mm., 9 18.5 mm.; ovipositor, 6 mm.

2 $, 2 9. Matamoras, Tamaulipas, Mexico, L. B. Couch ; Ringgold
Barracks at the lower end of the Rio Grande, C. A. Schott ; Carrigo
Springs, Texas, A. Wadgymar, through L. Bruner.

2. CEUTHOPHILUS ENSIFER.

Ceuthophilus ensifer Pack.!, Amer. Nat., xv. 882, pi. 7, figs. 4, 4ab
(1881); Id.!, Mem. Nat. Acad. Sc.,iv. 71-72, 83, figs. 17, 17ab (1888) ;
Blatchl., Proc. Ind. Acad. Sc., 1892, 153 (1894).

Body luteo-testaceous, heavily marked with blackish fuscous, which
broadly borders all the abdominal segments posteriorly and inferiorly,
as it does also (but more or less broken mediodorsally) the thoracic;
there are also two large subdorsal anterior blackish fuscous spots
on the meso- and metanotum, and the pronotum has a large T-shaped
blackish median spot heavily mapped out with a basal expansion and
which is cut by a mediodorsal luteous line, bordered on either side
posteriorly by one, anteriorly by two luteous dots, transversely aligned ;
the inferior border is broadly margined with black, leaving in the
middle of either side a large irregular luteous blotch more or less
streaked with fuscous ; the legs are castaneous, the hind femora with
fuscous scalariform markings and apically broadly annulate with
fuscous ; the apical half of tibiae and the tarsi pallid. The antennae
are about twice the length of the body and the legs long and slender.
The fore femora are not stouter than the middle femora, nearly twice
as long as the pronotum and somewhat more than half as long as the
hind femora, the inner carina with 3-4 spines of considerable size.
Middle femora with four spines on the front carina and on the hind
carina three besides a moderately long genicular spine. Hind femora
about as long as the body, somewhat less than twice as long as the
fore femora, slender, the apical third subequal, the inferior margin



SCUDDER. NORTH AMERICAN CEUTHOPHILI.

nearly straight, the whole fully four times as long as broad, the dark
portions of the surface, even of the inner side, with slight, equally
distributed raised points but none independent of them, the outer
carina ( 9 ) with delicate distant spinules especially beyond the middle,
the inner carina similarly armed, the intervening sulcus moderately
narrow. Hind tibiae considerably longer than the femora, slender,
armed beneath with a single preapical spine besides the apical pair ;
spurs rudely opposite, the basal well beyond the end of the proximal
third of the tibia, half as long again as the tibial depth, set at an
angle of 35 with the tibia and divaricating 90-100, their tips strongly
incurved ; inner middle calcaria considerably longer than the outer,
much more than twice as long as the others or as the spurs, but much
shorter than the first tarsal joint. Hind tarsi almost half as long as
the tibiae, the first joint as long as the rest together, the second almost
three times as long as the third and with it longer than the fourth.
Cerci tapering throughout, finely pointed, half as long again as the
femoral breadth. Ovipositor hardly three quarters as long as the fore
femora, stout in basal third, tapering in middle third, slender and sub-
equal in distal third, the apex produced and slightly upturned, the
inner valves with eight sharp but slight serrations.

Length of body, 9 19.5 mm. ; pronotum, 5.2 mm.; fore femora, 10
mm.; hind femora, 18.75 mm.; hind tibiae, 20.25 mm. ; ovipositor,
7 mm.

2 9 . Nickajack Cave, Tenn.

3. CEUTHOPHILUS STYGIUS.

Rhaphidophora stygia Scudd. !, Proc. Bost. Soc. Nat. Hist., viii. 9
(1861); Pack., Amer. Nat., v. 745 (1871).

Ceuthophilus stygius Scudd.!, Bost. Journ. Nat. Hist., vii. 438
(1862) ; Walk., Cat. Derm. Salt. Brit. Mus., i. 202 (1869); Pack.,
Guide Ins., 565 (1869) ; Riley, Stand. Nat. Hist., ii. 184 (1884) ;
Pack., Mem Nat. Acad. Sc., iv. 70-71, 83 (1888); Brunn., Monogr.
Stenop., 65 (1888); Blatchl., Proc. Ind. Acad. Sc., 1892, 148-149
(1894).

Ceuthophilus sloanii Pack. !, Ann. Rep. Peab. Acad. Sc., v. 93-94
(1873) ; Id. !, Mem. Nat. Acad. Sc., iv. 71, 83 (1888). Immature.

Body pale brown, the segments bordered posteriorly with dark
brown or black, becoming gradually paler toward the hinder part of
the body and dotted with pale spots ; the markings in general closely
resemble those of C. gracilipes, but the dark colors do not generally pre-
vail to so great an extent as in that species. The antennae, moderately
VOL. xxx. (N. s xxii.) 3



34 PROCEEDINGS OF THE AMERICAN ACADEMY.

coarse at the base, taper very gradually, and are of immense length,
being more than four times the length of the body. The legs are
very long and slender. Fore femora scarcely broader than middle
femora, very slender, three fourths as long again as the pronotum, yet
only about half as long as the hind femora, the inner carina armed
with 2-5 spines. Middle femora with 2-5 spines on the front cariua
and on the hinder 1-3 spines besides a not very long genicular spine.
Hind femora about as long as the body and twice the length of
the fore femora, not very stout, about four times as long as broad,
tapering gracefully, the apical third being subequal, the darker por-
tions covered with exceedingly fine and subdued raised points, the
outer carina with more or less slight distant serrations or triangular
denticulations occasionally developing as spines, especially on the
apical half, the inner carina with a few slighter distant similar serra-
tions closest in the middle half, the intervening sulcus not very broad.
Hind tibiae slender, straight in both sexes, fully a tenth longer than
the femora, armed beneath with 2-3 subapical spines besides the apical
pair; spurs vaguely opposite or subopposite, the basal within the
proximal fourth of the tibia, about half as long again as the tibial
depth, set at an angle of about 40 with the tibia and divaricating
about 90, their tips much incurved ; inner middle calcaria much
longer than the outer, more than twice as long as the others or as the
spurs and as long as the first joint of the tarsus, beset with short hairs,
as are also the upper calcaria. Hind tarsi fully two fifths the length
of the tibia, the first joint shorter than the rest combined, the second
twice as long as the third and with it shorter than the fourth. Cerci
rather long and slender, tapering to a fine point, fully half as long as
the hind femora. Ovipositor three fifths the length of the hind femora,
not greatly swollen at base, tapering gently in basal half, equal beyond
and not very narrow, the tip scarcely upturned and not produced, the
apex being nearly rectangular, the teeth of the inner valves feebly
crenate.

Length of body, $ 17 mm., 9 19.5 mm.; pronotum, 5.5 mm.,
9 5.3 mm.; fore femora, $ 9 9.75 mm.; hind femora, $ 19.5 mm.,
9 20.1 mm.; hind tibiae, $ 21.5 mm., 9 22.8 mm.; ovipositor,
13 mm.

1 <J, 3 9. Hickman's Cave, Ky., A. Hyatt ; cave in Crawford Co.,
Ind., W. P. Hay, through W. S. Blatchley. In the Museum of Com-
parative Zoology there are specimens taken at White's Cave, near
Mammoth Cave, Ky.; Fountain's Cave, A. S. Packard ; and One
Hundred Dome Cave, near Glasgow Junction, Ky., F. G. Sanborn,



SCUDDER. NORTH AMERICAN CEUTHOPHILI. 35

II. Garman. Packard, in his Memoir on Cave Animals, reports it
also from Little and Great Wyandotte Caves, a cave in Washington
Co., and Georgetown, Floyd Co., all in Indiana, and also from the
following caves in Kentucky : Diamond Cave, near Mammoth Cave,
John and Fred Field Cave, near Dismal Creek, Bee Spring and
Laurel Caves, Carter Co. According to Brunner, specimens seen by
him come from Texas. C. sloanii was described from caves in Ken-
tucky and Bradford, Ind.

4. CEUTHOPHILUS GRACILIPES.

Phalangopsis gracilipes Hald., Proc. Amer. Assoc. Adv. Sc., ii. 346
(1850) ; Walk., Cat. Derm. Salt. Brit. Mas., i. 116 (1869).

Rhaphidophora gracilipes Scudd. !, Proc. Bost. Soc. Nat. Hist., viii. 7
(1861).

Ceuthophilus gracilipes Scudd. !, Bost. Journ. Nat. Hist., vii. 439
(1862) ; Walk., Cat. Derm. Salt. Brit. Mus., i. 202 (1869) ; Scudd. !,
Rep. U. S. Geol. Surv. Nebr., 249 (1872) ; Brunn., Monogr. Stenop.,
62-63 (1888) ; Smith, Cat. Ins. N. J., 409 (1890) ; McNeill, Psyche,
vi. 27 (1891).

Ground color of body varying from luteous to dark castaneous, very
heavily marked with blackish fuscous, so that the latter is often or
perhaps generally the prevailing tint; the darker colors prevail always
on the hinder half of all the segments but the pronotum and some-
times, especially in young specimens, to such an extent that all parts
behind the pronotum are blackish fuscous, dotted with luteous ; in the
lightest colored specimens, the dark coloring prevails on the pronotum
along the anterior and especially the posterior margins, and is generally
present in a subdorsal posterior tongue emitted on either side from the
anterior margin ; the anterior edge of the dark posterior markings of
each segment, especially in the front portion of the body, is exceed-
ingly irregular and broken, and the lightest parts are more or less and
irregularly clouded with fuscous ; the femora are usually of the pre-
vailing tint of the body, but, even when the body is dark, are some-
times luteous throughout as the tibia? and tarsi always are, except for
occasional infuscation of the former at base ; the outer sides of the
hind femora have the characteristic markings of the genus more or less
distinct. Antennse moderately coarse at base, tapering with great
regularity, 3-4 times the length of the body. Legs very long and
slender. Fore femora no stouter than the middle femora, more than
half as long again as the pronotum, especially in the , distinctly less
than half as long as the fore femora, the inner carina usually with 2-3



36 PROCEEDINGS OF THE AMERICAN ACADEMY.

spines, often rather long in old individuals, especially the distal ones.
Middle femora with 3-4 spines on either carina, the hind carina
with a not very long genicular spine. Hind femora as long as or
longer than the body, considerably more than twice the length of the
fore femora, very stout at base, but gradually diminishing in stout-
ness so that the distal third is slender and subequal and the whole
less than three and a half times longer than broad, the surface mostly
glabrous, but on the darker portions above beset not very sparsely
with feeble raised points, the outer carina with about thirteen distant un-
equal rather coarse spines, the longest shorter than the tibial spurs ( $ )
or almost unarmed (9), the inner carina much less elevated than
the outer, with equal abortive denticulations numerous in the , in-
frequent and equidistant in the 9 , the intervening sulcus moderately
broad. Hind tibiae rather slender, straight or in old male specimens
gently subarcuate or slightly waved in the proximal third, nearly a
sixth longer than the femora, armed beneath usually with two aligned
preapical spines besides the apical pair ; spurs subopposite, the basal
pair near end of proximal fourth of tibia, less than half as long
again as the tibial depth, set at an angle of about 40 with the tibia
and usually divaricating about 60, but sometimes to as much as
twice that, incurved especially at tip; inner middle calcaria much
longer than the outer, more than twice as long as the other calcaria,
about twice the length of the spurs, but much shorter than the first
tarsal joint. Hind tarsi much less than half as long as the tibia, the
first not so long as the rest together, the second nearly three times as
long as the third and with tt fully as long as the fourth. Cerci stout at
base, tapering throughout, nearly a third as long as the hind femora.
Ovipositor with the basal third rather stout, beyond equal and rather
slender, nearly three fourths the length of the hind femora, the arma-
ture of inner valves acicular, arcuate, elongate.

Length of body, $ 19 mm., 9 23 mm. ; antennae, $ (est.) 75 mm. ;
pronotum, g 5.75 mm., 9 6-75 mm. ; fore femora, $ 10 mm., 9 10-6
mm.; hind femora, $ 21.5mm., 9 22 mm.; hind tibia;, $ 24.75mm.,
9 25 mm.; ovipositor, 15.5 mm.

22 ,11 9. Maryland, New Jersey, P. R. Uhler ; Ithaca, N. Y.,
Comstock; Blockton, Florida (C. Caule, Jr.), J. H. Comstock;
Southern Illinois, P. R. Uhler; Illinois, Walsh, Webster; Northern
Illinois, R. Kennicott ; Minnesota ; Red River, Manitoba, D. Gunn ;
Nebraska City, Nebr., F. V. Hayden. It has also been reported from
Pennsylvania by Haldeman, and from Georgia and Colorado by
Brunner.



SCUDDER. NORTH AMERICAN CEUTHOPHILI. 37

5. CEUTHOPHILUS LATEBRICOLA, sp. nov.

Blackish fuscous, sometimes almost piceous, glabrous, profusely spot-
ted with luteous or rufo-luteous and more or less blotched with the same
on the thoracic segments, though nowhere in large masses, but, except
the luteous bordering of the inferior margins, more as if through the
confluence of minute spots. There is always a narrow mediodorsal line
or stripe on the thoracic segments with slight expansions anteriorly and
posteriorly on the pronotum ; legs luteous with heavy blackish infusca-
tions at the femoral tips, the hind femora heavily marked with fuscous
scalariform markings. The antennae are slender and about three
times as long as the body, and the legs moderately long and slender.
Fore femora no stouter than the middle femora, considerably less than
half as long as the hind femora, a little more ( <J) or a little less ( 9 )
than a third longer than the pronotum, the inner carina with a long
preapical spine and sometimes another much smaller. Middle femora
with 2-4 spines on the front carina and on the hind carina with 1-2
spines besides a long genicular spine. Hind femora about as long as
the body, considerably more than twice as long as the fore femora,
moderately stout, in the male somewhat strongly constricted before the
genicular lobes, about three and a quarter times longer than broad (<),
the surface with only a few raised points along the upper edge interi-
orly, the outer carina considerably elevated before the constriction,
armed with 6-9 unequal teeth, the largest tumid at base and not so
long as the tibial spurs (^) or with 23 feeble serrations (9), the
inner carina with distant feeble minute spinules (9)? the intervening
sulcus slender. Hind tibiae sinuate in the proximal half ( <?) or straight
throughout (9), considerably longer than the femora (scarcely longer
in Eastern examples), armed beneath with a single subapical spine
besides the apical pair ; spurs subopposite, the basal just before the
end of the proximal third of the tibia, about half as long again as the
tibial depth, set at an angle of 45 with the tibia and divaricating
100, their apical third incurved ; inner middle calcaria very much
longer than the outer, almost twice as long as the others or as the
spurs and nearly as long as the first joint of the tarsi. Hind tarsi
not a third as long as the tibia?, the first joint longer than the rest
together, the second fully twice as long as the third and with it at
least as long as the fourth. Cerci rather stout, rapidly tapering,
a little longer than half the femoral width. Abdomen roundly trun-
cate in the male. Ovipositor straight, two thirds as long as the hind
femora, gently tapering in basal third, beyond equal and slender, the



38 PROCEEDINGS OF THE AMERICAN ACADEMY.

tip considerably upcurved and very acuminate, the teeth of the inner
valves aculeate.

Length of body, $ 15 mm., 9 15.5 mm.; antennae, $ 52 mm.;
pronotum, $ 5 mm., 9 4.75 mm. ; fore femora, ,6.9 mm., 9 6 mm. ;
hind femora, $ 15.5 mm., 9 13.5 mm. ; hind tibiae, $ 17.5 mm.,
9 14.5 mm.; ovipositor, 9 mm.

4 (, 6 9- Lexington and Tyrone, Ky. (S. Garman) ; Washington,
D. C., Wright; Centre Co., Peun., Shaler ; and Petroleum, Eitchie
Co., W. Va. (Mus. Comp. Zool.).

6. CEUTHOPHILUS GRANDIS, sp. nov.

Body dark luteo-castaneous very heavily marked with blackish or
blackish fuscous, which is heaviest at the posterior margins of all the
segments, along the front margin of the pronotum, and in a stripe
bordering the broad mediodorsal rufo-luteous stripe of the pronotum ;
the anterior lower angle of the pronotum, and to some extent that
of the other thoracic segments, are dull luteous, merging with the
luteous of the central portion of the descending lobes ; on the abdomen
the darker colors prevail above to such a degree that the luteous
appears to be maculate on a dark ground ; antennas and legs luteo-
testaceous, more or less though generally feebly infuscate, the hind
femora with fuscous scalariform markings, apically broadly marked
with fuscous but preceded by a broad more or less clearly marked
luteous annulation. The antennae are slender except near the base
and fully four times as long as the body, the legs very long and
slender. Fore femora scarcely stouter than the middle femora, barely
half as long as the hind femora, two thirds (<J) or three fifths ( 9 ) as
long again as the pronotum, the inner carina with 3-4 spines none of
them very long nor very unequal. Middle femora similarly armed
on the front carina and on the hind carina 3-4 spines besides the
moderately long genicular spine. Hind femora as long as the body,
very little more than twice as long as the fore femora, pretty stout at
base, but with the slender portion much produced, the apical third or


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